The geodynamic evolution of the East Junggar is examined by means of satellite imaging and field-based structural studies, U-Pb zircon geochronology and analysis of potential field geophysical data in the Yemaquan arc and the Dulate back-arc systems. The northern Yemaquan arc shows a pervasive WNW-ESE steep S₁ foliation that is related to the exhumation of Armantai ophiolitic mélange in an F₁ antiformal structure. The bedding of the Dulate sequences is folded by N-S-trending F₁ upright folds that are preserved in low strain domains. The timing of D₁ is estimated between 310 and 280 Ma. During D₂, previously folded Dulate sequences were orthogonally refolded by E-W-trending F₂ upright folds, resulting in Type-1 basin and dome interference pattern and pervasive E-W trending S₂ cleavage zones. The age of D₂ is constrained to be 270-250 Ma based on the dating of syn-tectonic pegmatites and deposition of syn-orogenic sedimentary rocks. The boundary between the Yemaquan arc and Dulate back-arc basin experienced reactivation through D₂ dextral transpressive shear zones. The D₁ fabrics are the consequence of the closure of the Dulate back-arc basin due to the advancing mode of Kalamaili subduction. Almost orthogonal Permian D₂ fabrics were generated by the N-S shortening of the East Junggar and the northward movement of the Junggar Block indenter. This D₂ deformation was associated with the anticlockwise rotation of the southern limb of the Mongolian Orocline, the scissor-like closure of the northerly Mongol-Okhotsk Ocean and the collision of the Mongolian and the Tarim-North China craton collages.